| Name | Jurisdiction of Incorporation | |||||||
| Apeiron Biologics GmbH | Austria | |||||||
| CyDex Pharmaceuticals, Inc. | Delaware | |||||||
| LHNC, Inc. | Delaware | |||||||
| Ligand Holdings UK Ltd. | England and Wales | |||||||
| Ligand UK Development Limited | England and Wales | |||||||
| Ligand UK Limited | England and Wales | |||||||
| Metabasis Therapeutics, Inc. | Delaware | |||||||
| Neurogen Corporation | Delaware | |||||||
| Pelthos Therapeutics Inc. | Delaware | |||||||
| Pfenex Inc. | Delaware | |||||||
| Pharmacopeia, LLC | Delaware | |||||||
| Seragen, Inc. | Delaware | |||||||
| Vernalis Therapeutics, Inc. | Delaware | |||||||