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S-3 EX-FILING FEES 0001768224 N/A N/A 0001768224 1 2025-12-17 2025-12-17 0001768224 2 2025-12-17 2025-12-17 0001768224 3 2025-12-17 2025-12-17 0001768224 4 2025-12-17 2025-12-17 0001768224 5 2025-12-17 2025-12-17 0001768224 6 2025-12-17 2025-12-17 0001768224 7 2025-12-17 2025-12-17 0001768224 8 2025-12-17 2025-12-17 0001768224 9 2025-12-17 2025-12-17 0001768224 2025-12-17 2025-12-17 iso4217:USD xbrli:pure xbrli:shares

Ex-Filing Fees

CALCULATION OF FILING FEE TABLES

S-3

Arcturus Therapeutics Holdings Inc.

Table 1: Newly Registered and Carry Forward Securities

                                                             
Line Item Type   Security Type   Security Class Title   Notes   Fee Calculation
or Carry Forward Rule		   Amount Registered   Proposed Maximum Offering
Price Per Unit		   Maximum Aggregate Offering Price   Fee Rate     Amount of Registration Fee   Carry Forward Form Type   Carry Forward File Number   Carry Forward Initial Effective Date   Filing Fee Previously Paid in Connection with
Unsold Securities
to be
Carried Forward
                                                             
Newly Registered Securities
Fees to be Paid   Equity   Common Stock, $0.001 par value per share (1)        457(o)       $     $         $                 $  
Fees to be Paid   Equity   Preferred Stock, $0.001 par value per share (1)        457(o)                                            
Fees to be Paid   Debt   Debt Securities        457(o)                                            
Fees to be Paid   Other   Warrants        457(o)                                            
Fees to be Paid   Other   Units        457(o)                                            
Fees to be Paid   Other   Rights        457(o)                                            
Fees to be Paid   Unallocated (Universal) Shelf        (1)   457(o)               12,234,681.00   0.0001381     1,670.00                  
Fees Previously Paid                                                             
Carry Forward Securities
Carry Forward Securities   Unallocated (Universal) Shelf   Unallocated (Universal) Shelf    (2)   415(a)(6)       $     $ 487,765,319.00       $     S-3   333-269003   12/30/2022   $ 53,752.00
                                                             
Total Offering Amounts:   $ 500,000,000.00         55,422.00                  
Total Fees Previously Paid:               53,752.00                  
Total Fee Offsets:               0.00                  
Net Fee Due:             $ 1,670.00                  

__________________________________________
Offering Note(s)

(1) (i) An unspecified number of securities or aggregate principal amount, as applicable, is being registered as may from time to time be offered at unspecified prices and, in addition, an unspecified number of additional shares of common stock is being registered as may be issued from time to time upon conversion of any debt securities that are convertible into common stock or pursuant to any anti-dilution adjustments with respect to any such convertible debt securities.

(ii) Includes rights to acquire common stock or preferred stock of the Company under any shareholder rights plan then in effect, if applicable under the terms of any such plan.

(iii) Estimated solely for the purpose of calculating the registration fee. No separate consideration will be received for shares of common stock that are issued upon conversion of debt securities or preferred stock or upon exercise of common stock warrants registered hereunder. The aggregate maximum offering price of all securities issued pursuant to this registration statement will not exceed $500,000,000.

(iv) The registration fee has been calculated pursuant to Rule 457(o) under the Securities Act of 1933, as amended.
(2) (2) Pursuant to Rule 415(a)(6), the securities being registered hereunder include $487,765,319 of unsold securities which remain unsold as of the date hereof (the “Unsold Securities”) previously registered by the registrant’s registration on Form S-3 (File No. 333-269003) which was initially filed with the Securities and Exchange Commission on December 23, 2022 and declared effective on December 30, 2022 (the “Prior Registration Statement”). The aggregate filing fee paid in connection with such Unsold Securities was $53,752 (based on the filing fee rate in effect at the time of the filing of the Prior Registration Statement) and will continue to be applied to the Unsold Securities that are being carried forward to this registration statement. No additional filing fee is due with respect to the Unsold Securities carried forward in this registration statement. To the extent that, after the filing date hereof and prior to the effectiveness of this registration statement, the Registrant sells any Unsold Securities pursuant to the Prior Registration Statement, the Registrant will identify in a pre-effective amendment to this registration statement the updated amount of Unsold Securities from the Prior Registration Statement to be included in this registration statement pursuant to Rule 415(a)(6) and the updated amount of new securities to be registered on this registration statement, if any. Pursuant to Rule 415(a)(6), the offering of securities under the Prior Registration Statement will be deemed terminated as of the date of effectiveness of this registration statement.